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Hola Rent a Car claims to be the safest rent-a-car company in the country. They even have a satellite system to unlock your car door anywhere in the country should you accidentally lock your keys in the car. They have main rental offices in San José, Guanacaste, and in some hotels such as the Marriott Courtyard and Costa Rica Marriott close to the airport. Their web page offers more information about where you can pick up or drop off rental cars. On their web site is a map indicating where you can find their offices.
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One of the best guide books about Costa Rica we have found